Up‐regulation of <scp>PR</scp> 1 and less disruption of hormone and sucrose metabolism in roots is associated with lower susceptibility to ‘ Candidatus Liberibacter asiaticus’


Publication

Citation
Wu et al. (2018). Plant Pathology 67 (6)
Names (1)
Subjects
Agronomy and Crop Science Genetics Horticulture Plant Science
Abstract
Huanglongbing ( HLB ), caused by ‘ Candidatus Liberibacter asiaticus’ (Las), is a devastating disease of citrus trees in Florida. Previous work showed that the rootstock cultivar Cleopatra mandarin ( Citrus reticulata ) has a higher population of Las in roots than Swingle citrumelo ( C. paradisi  ×  Poncirus trifoliata ). Las reduced fibrous root biomass and sucrose content in Cleopatra mandarin more than in Swingle citrumelo. To understand the mechanisms for susceptibility to Las infection, sucrose and hormone metabolism status were evaluated in Cleopatra mandarin and Swingle citrumelo. In fibrous roots of Cleopatra mandarin, higher expression of genes related to sucrose cleavage was consistent with lower sucrose content compared to noninoculated seedlings at 5 weeks post‐root trimming (wpt). In fibrous roots of Swingle citrumelo, both sucrose content and gene expression related to sucrose cleavage were less disrupted by Las infection compared to Cleopatra mandarin at 5 wpt. Genes associated with salicylic acid ( SA ), ethylene ( ET ) and abscisic acid ( ABA ) synthesis, and ABA signalling, phospholipases D ( PLD ), and phospholipase A2 ( PLA 2) were activated by Las infection at 5 wpt in Cleopatra mandarin. Expression of downstream effectors of SA , i.e. NPR 1, WRKY 70 and PR 1, did not change in Cleopatra mandarin, suggesting inhibition of the response to SA by the elevation of ABA , ET and PLD . In contrast, the up‐regulation of PR 1, lower response of sucrose metabolism genes and down‐regulation of biosynthesis of phytohormones indicates that Swingle citrumelo activates a more effective defence against this biotrophic pathogen than Cleopatra mandarin.
Authors
Wu, J.; Alférez, F. M.; Johnson, E. G.; Graham, J. H.
Publication date
2018-08-01
DOI
10.1111/ppa.12838 

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