Cell is gram-negative, aerobic, non-motile, and rod-shaped (0.7–2.0-μm long and 0.3–0.7-μm wide). Colonies are circular with regular margins, convex, milky, and 2–3 mm in diameter after incubation for 48 h on YMA at 30 °C. Grows in the presence of 0–2% (w/v) NaCl (optimum, 0% NaCl) at pH 6.0–9.0 (optimum, 7.0–8.0) and at 15–37 °C (optimum, 30 °C). Positive for nitrate reduction, and catalase and oxidase. Eesculin is hydrolyzed and gelatin is weakly hydrolyzed. Starch and casein are not hydrolyzed. Can assimilate mannose, N-acetyl glucosamine, maltose, and citric acid. Negative for H2S, urease, and indole production and Voges–Proskauer reaction. In the API ZYM system, positive for alkaline phosphatase, esterase (C4), esterase lipase (C8), leucine arylamidase, acid phosphatase, naphthol-AS-BI-phosphohydrolase, and β-glucosidase. The following carbon sources are utilized: dextrin, N-acetyl-d-glucosamine, N-Acetyl-β-d-mannosamine, α-d-glucose, d-mannose, d-fructose, d-galactose, d-fucose, l-fucose, l-rhamnose, d-mannitol, d-arabitol, myo-inositol, glycerol, glycyl-l-proline, l-alanine, l-aspartic acid, l-glutamic acid, d-galacturonic acid, d-gluconic acid, d-glucuronic acid, glucuronamide, α-keto-glutaric acid, d-malic acid, l-malic acid, bromo-succinic acid, Tween 40, β-hydroxy-d, l-butyric acid, acetoacetic acid, and acetic acid. The major cellular fatty acids are summed feature 8 (C18:1ω7c and/or C18:1ω6c) and C16:0. The draft genome is 4.75 Mb in size and the common genes required for legume nodulation, nodACD and nifH, are absent from the genome. The G + C content is 60.0%.